Lasioglossum (Dialictus) gaudiale

Lasioglossum (Dialictus) gaudiale (Sandhouse, 1924) Figs 24–26, 76E, 91B, 94A, 103A, 104A, 114A Halictus (Chloralictus) gaudialis Sandhouse, 1924: 23 (holotype, ♂, deposited in USNM, type no. 26418, examined). Halictus (Chloralictus) gaudialis – Michener 1936: 283 (relationship to helianthi). Lasioglossum (Chloralictus) tegulariforme – Michener 1951: 1118 (in part, synonymy, catalogue). Dialictus tegulariformis – Hurd 1979: 1972 (in part, catalogue). — Moure & Hurd 1987: 134 (in part, catalogue). Lasioglossum (Dialictus) gaudiale – Gardner & Gibbs 2022: 28 (resurrection from synonymy). Diagnosis Females of L. gaudiale have the tegula relatively small (reaching and sometimes slightly exceeding posterior margin of mesoscutum in dorsal view), with inner posterior margin strongly concave and a small rounded posterior projection, and usually densely punctate (IS ≤ 1 PD, rarely up to 2 PD); T1 anterior slope polished and mirror-smooth (rarely with very weak microsculpture basally); gena about as wide as eye in lateral view; metapostnotum usually shiny basally with strong subparallel rugae not reaching the posterior margin (sometimes dull, with rugae anastomosing, or rugae reaching posterior margin, but never all at once); mesoscutum with no tomentum; T3 without a subapical band of tomentum; and metasoma usually black to brown (rarely red-orange). In many cases, the scutellum is significantly more coarsely and sparsely punctate than the mesoscutum (IS = 1–2 PD), but this is unreliable as a primary diagnostic character. Females of L. gaudiale are most similar to those of L. angelicum sp. nov., L. eremum sp. nov., L. gloriosum sp. nov., L. indagator sp. nov., and L. pseudotegulare, all of which (except L. indagator) have the gena narrower than eye in lateral view and scutellum usually finely and densely punctate (IS <1 PD). In addition, females of L. eremum sp. nov. have the metapostnotum with anastomosing rugae reaching the posterior margin and T1 anterior slope coriarious. Females of L. gloriosum have the T1 anterior slope coriarious, mesoscutum lateral and posterior margins with dense tomentum, metapostnotum rugae usually reaching the posterior margin, and metasoma red-orange. Females of L. indagator have the tegula sparsely punctate (IS = 1–3 PD) and inner posterior margin weakly concave. Females of L. pseudotegulare have the metapostnotum entirely dull with fine rugae and tegula very large (exceeding posterior margin of mesoscutum in dorsal view) and more densely punctate (IS <1 PD). Females of L. angelicum sp. nov. have a similarly large tegula. Males of L. gaudiale have the tegula relatively small (reaching or slightly exceeding posterior margin of mesoscutum in dorsal view), with inner posterior margin strongly concave and a narrow posterior point, and usually densely punctate (IS <1 PD); face relatively narrow (length/width ratio usually>0.83) with dense tomentum below the upper ocular tangent (often surrounding the median ocellus); metapostnotum usually shiny with strong subparallel rugae not reaching posterior margin; mesoscutum usually densely punctate (IS ≤ 1 PD) and fringed with tomentum; mesepisternum somewhat coarsely punctate with distinct interspaces (IS ≤1 PD); preëpisternum and metepisternum with sparse appressed tomentum; T2–T3 with abundant basolateral tomentum; and head and mesosoma integument usually deep blue. Males of L. gaudiale are most similar to those of L. gloriosum sp. nov. and L. indagator sp. nov. Both of these species have the face broad (length/width ratio ≤ 0.83) and preëpisternum and metepisternum with more abundant tomentum. In addition, males of L. gloriosum have the mesepisternum very densely punctate (IS <1 PD); metapostnotum usually with anastomosing rugae reaching posterior margin; and integument usually greener. Males of L. indagator have the metapostnotum with very short parallel rugae not reaching the posterior margin; tegula inner posterior margin more weakly concave, forming a blunt angle posteriorly, and usually moderately sparsely punctate laterally (IS = 1–2 PD); and mesoscutum usually slightly more sparsely punctate (IS = 1–2 PD). Etymology Sandhouse (1924) formed the specific epithet ʻ gaudialis ʼ from the Latin noun ʻ gaudium ʼ (joy) and the adjectival suffix ʻ -alis ʼ (state or condition of), together meaning ʻjoyfulʼ. Material examined Holotype UNITED STATES – California • ♂; La Jolla; [32.83° N, 117.27° W]; August [day/year unknown]; Cockerell leg.; USNM 26418. Other material MEXICO – Aguascalientes • 1 ♀; 14 mi. W of Aguascalientes; [21.88° N, 102.51° W]; 10 Jul. 1983; P.W. Kovarik, Harrison, J.C. Schaffner leg.; TAMU-ENTO X1416976. – Baja California • 1 ♀; “ Gonzales Bay ” [Bahía de San Luis Gonzaga]; [29.79° N, 114.4° W]; 29 Apr. 1921; E.P. Van Duzee leg.; UCMC. – Baja California Sur • 1 ♀; 2 miles north of Todos Santos; 23.4687° N, 110.2399° W; 20 Oct. 2016; Paul A. Rude leg.; EMEC • 1 ♀; ibid.; EMEC • 1 ♀; El Pedrito beach, 3 mi. S of Todos Santos; 23.3941° N, 110.2124° W; 24 Sep. 2016; Paul A. Rude leg.; EMEC • 1 ♂; La Paz; [24.14° N, 110.31° W]; 3 Jun. 1921; E.P. Van Duzee leg.; CAS • 1 ♀; La Paz; 24.1446° N, 110.31° W; 20 Feb. 1974; George E. Bohart leg.; BBSL1101069 • 2 ♀♀, 1 ♂; La Paz; 24.1742° N, 110.304° W; 5 Sep. 2016; Paul A. Rude leg.; EMEC • 1 ♀; La Paz, 2 miles south; [24.08° N, 110.31° W]; 31 Jul. 1966; E.G. Linsley, J.M. Linsley leg.; EMEC • 1 ♀; Monserrate Island, Gulf California; [25.68° N, 111.03° W]; 13 Jun. 1921; E.P. Van Duzee leg.; CAS • 1 ♀; Todos Santos; 23.4587° N, 110.2335° W; 8–12 Oct. 2018; Paul A. Rude leg.; EMEC. – Chihuahua • 1 ♀; Juarez; 31.7353° N, 106.4721° W; 22 Jul. 1971; D.W. Davis leg.; BBSL1101081 • 2 ♀♀; Municipio Buenaventura, 9 mi. N of Flores Magón; [30.07° N, 106.96° W]; 29 Jun. 1981; NMSU. – Coahuila • 1 ♂; Boquillas; 29.1883° N, 102.9374° W; 19 Jun. 1992; Robert L. Minckley leg.; ex Haplopappus; SEMC. – Sonora • 1 ♂; 10 mi. S of Guaymas; 27.9102° N, 110.7259° W; 7 Oct. 1968; George E. Bohart leg.; ex Tecoma stans; BBSL1100989 • 5 ♀♀; Rancho Puerta Blanca; 31.3119° N, 109.1133° W; 25 May 2008; Robert L. Minckley leg.; RLM SBV083922, SBV083928, SBV083942, SBV083968, SBV083996 • 1 ♀; Rancho San Bernardino; 31.3136° N, 109.2547° W; 29 Aug. 2001; Robert L. Minckley leg.; RLM SBV137953 • 1 ♀; ibid.; 13 Aug. 2002; A. Romero leg.; RLM SBV043987 • 1 ♂; Rancho San Bernardino; 31.3139° N, 109.2503° W; 3 Jun. 2002; Robert L. Minckley leg.; RLM SBV016827 • 1 ♀; Rancho San Bernardino; 31.316° N, 109.2576° W; 6 Jun. 2004; Robert L. Minckley leg.; ex Baileya multiradiata; RLM SBV025444 • 1 ♀; ibid.; 10 Jun. 2004; N. de la Torre leg.; RLM SBV025674 • 1 ♀; Rancho San Bernardino; 31.3169° N, 109.2592° W; 6 Jun. 2004; N. de la Torre leg.; RLM SBV025607 • 1 ♀; ibid.; 15 Jun. 2004; N. de la Torre leg.; RLM SBV054797 • 1 ♀; Rancho San Bernardino; 31.3179° N, 109.2602° W; 15 Jun. 2004; N. de la Torre leg.; ex Baileya multiradiata; RLM SBV054768 • 3 ♀♀; Rancho San Bernardino; 31.3195° N, 109.2701° W; 18 Aug. 2003; A. Nelson leg.; RLM SBV035698, SBV035715, SBV035722 • 1 ♀; ibid.; 19 Aug. 2003; A. Nelson leg.; RLM SBV036076 • 1 ♀; Rancho San Bernardino; 31.3195° N, 109.2703° W; 8 May 2001; Robert L. Minckley leg.; RLM SBV019590 • 1 ♂; ibid.; 8 May 2001; Robert L. Minckley leg.; RLM SBV019591 • 1 ♀; Rancho San Bernardino; 31.3196° N, 109.2693° W; 18 Aug. 2003; A. Nelson leg.; RLM SBV035622 • 12 ♀♀; Rancho San Bernardino; 31.32° N, 109.2685° W; 18 Aug. 2003; A. Nelson leg.; RLM SBV035642, SBV035649, SBV035650, SBV035651, SBV035652, SBV035653, SBV035659, SBV035660, SBV035661, SBV035665, SBV035666, SBV035670 • 1 ♀; ibid.; 19 Aug. 2003; A. Nelson leg.; RLM SBV035755 • 6 ♀♀; Rancho San Bernardino; 31.3204° N, 109.2681° W; 2 Sep. 2003; A. Romero leg.; RLM SBV036595, SBV036596, SBV036606, SBV036613, SBV036631, SBV036636 • 1 ♀; Rancho San Bernardino; 31.3319° N, 109.2567° W; 11 Jun. 2004; A. Romero leg.; RLM SBV025922. – Unknown • 1 ♀; San Luis [exact location unknown]; 30 May 1955; G.D. Butler leg.; ex Heliotrope; UAIC. UNITED STATES – Arizona • 3 ♀♀; Cochise Co., 10 mi. E of Douglas; [31.35° N, 109.35° W]; 22 Aug. 1994; Jerome G. Rozen, John S. Ascher leg.; AMNH • 1 ♀; Cochise Co., 2 mi. W of Fort Huachuca; [31.55° N, 110.38° W]; 28 Jun. 1983; W.J. Pulawski leg.; CAS CASENT 8416787 • 1 ♂; Cochise Co., Benson; [31.97° N, 110.29° W]; 12 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; SEMC • 1 ♀; Cochise Co., Portal; 31.8833° N, 109.2055° W; 20 Jul. 2017; S. L. Buchmann leg.; WRME 518956 • 1 ♀; Cochise Co., Rancho San Bernardino; 31.3139° N, 109.2503° W; 16 Aug. 2000; K. Toal leg.; RLM SBV002658 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge; 31.3369° N, 109.2436° W; 27Aug. 2001; Robert L. Minckley leg.; RLM SBV137736 • 1 ♀; Maricopa Co., Marquahala; [33.8° N, 113.3° W]; 15 Sep. 1979; J. Haworth, G.D. Butler leg.; UAIC • 10 ♀♀; Maricopa Co., Phoenix, Cotton Research Center; [33.404° N, 111.988° W]; 20–21 Sep. 1979; G.D. Butler leg.; ex cotton; UAIC • 1 ♀; Maricopa Co., Rainbow Valley; [33.2° N, 112.4° W]; 7 May 1980; J. Gillespie leg.; CUIC • 1 ♀; Mohave Co., “L. Havaska” [Lake Havasu] City; [34.48° N, 114.32° W]; 2 Apr. 1973; George E. Bohart leg.; BBSL700789 • 1 ♀; Mohave Co., 0.87 air km ESE of Beaver Dam; 36.8972° N, 113.9242° W; 10 May 2014; M.C. Orr leg.; ex Tamarix ramosissima; BBSL907904 • 1 ♀; Mohave Co., 10 mi. S of Lake Havasu City; [34.38° N, 114.2° W]; 14 Mar. 1972; P.F. Torchio, B. Apperson leg.; ex Phacelia; BBSL700792 • 2 ♀♀; Mohave Co., 2 mi. N of “Topac” [Topock]; [34.81° N, 114.5° W]; 1 May 1983; P.F. Torchio leg.; ex Nama; BBSL700787, BBSL700788 • 1 ♀; Mohave Co., Topock; [34.77° N, 114.49° W]; 15 Mar. 1972; P.F. Torchio, B. Apperson leg.; ex Cercidium; BBSL700794 • 1 ♂; Pima Co., 2 km NE of Oracle; 32.626° N, 110.7362° W; elev. 1240 m; 20–24 May 2013; M.E. Irwin leg.; BBSL FDP152841 • 1 ♂; Pima Co., Tucson; [32.2° N, 110.9° W]; 2 Jul. 1991; W. T. Wcislo leg.; SEMC • 1 ♀; Santa Cruz Co., near E

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