Anorus piceus LeConte 1859

Anorus piceus LeConte, 1859 (Figs. 5–7) Anorus piceus LeConte 1859a: 87. Type locality: San Diego, California. Anorus piceous [lapsus calami]: Blaisdell 1934: 320. Diagnosis. As in A. parvicollis, males of this species (Fig. 5) are readily separated from Dascillus by the prosternal process forming a short denticle near the anterior coxal margin, resulting in the procoxal cavities being contiguous externally (prosternal process completely separating procoxae in Dascillus). It may be separated from A. parvicollis by the dense, recumbent setae present throughout the dorsal surface, including the scutellar shield, and lack of long erect setae, though many specimens have a pelage of suberect setae on the elytra (A. parvicollis bearing sparse, long, mostly erect setae). The pronotum is distinctly narrowest across the anterior angles, then the pronotum gradually widens posteriorly for two-thirds of the length of pronotum, then narrows slightly to the posterior angles (Fig. 5A). The elytron is more similar to that of Dascillus than to that of A. parvicollis, both in disposition of setae and the presence of a sulcus along the lateral margin extending from behind the humerus nearly to the apical angle. The mandibles are more elongate in this species than A. parvicollis, the incisor edge is longitudinally straight anterior to the labrum, and there are two retinacula (proximal one longer and more acute) before the apex. Females are easily recognized by the disposition of the elytra, which extend about a third to a half of the distance to the tip of the abdomen (Fig. 6; see also Blaisdell 1934: fig. 3). Redescription of Male. Body parallel-sided, moderately slender, 8–14 mm (Fig. 5A). Body setae of two types, most fairly short and recumbent; elytra, especially laterally, and appendages also bearing scattered moderately long, bristling setae. Antennomeres 3–10 weakly serrate, at least 2 times longer than wide. Left mandible bearing two strong retinacula, lacking basal angulation. Right mandible with two retinacula, basal retinaculum very strong and finger-like, more apical retinaculum blunt, appearing as a lobe on the incisor edge near the base of the apical tooth and often nearly absent. Epicranial suture faintly traceable. Lateral sides of pronotum weakly explanate, lateral carina always distinct to anterior margin, posterior margin weakly but distinctly bisinuate; pronotum widest in posterior third, anterior margin always narrower than posterior margin. Hypomeron with distinct but short postcoxal process. Prosternal process acute apically, lateral margins converging near middle of procoxae, continued as thin vertical lamina often concealed by procoxae (as in Fig. 6B); mesoventrite with abrupt mesal elevation projecting anteriorly, bearing small procoxal rests. Elytra attaining the apex of abdomen, with weakly punctate striate. Hind wing exceeding apex of abdomen; radial cell completely closed proximally; wedge cell medium-sized, separated from posterior edge by distance about half length of cell; apical field about one-third length of wing. All tibiae lacking stout spinules along outer margin. All legs with tarsomeres 1–4 bearing fleshy bilobed pads beneath, tarsomere 1 with relatively small lobes. Terminalia with tergite IX (Fig. 5G) with very broad and shallow V-shaped emargination along posterior margin; tergite X (Fig. 5G) evenly broadly rounded apically; sternite IX (Fig. 5F) rounded apically; phallobase of aedeagus (Figs. 5B–D) weakly emarginate to rounded basally, slightly broader than long; parameres narrow, much longer than phallobase, gradually narrowing from base to apex, lateral margin slightly concave, recurved apically, bearing very few scattered, fine setae in apical half; dorsal lobe of penis comparatively narrow, measured from anterior margin of basal strut to posterior apex approximately as long as paramere, variably narrowed apically, apex evenly to narrowly rounded and never appearing cleft; ventral lobe of penis nearly as long as dorsal lobe, curved dorsally around middle, then curved ventrally near apex, apex narrowly rounded. Redescription of Female. Resembling male in most characters except: Body stouter (Fig. 6), wider in apical half. Elytra shorter than abdomen, about 1.5–2× as long as pronotum, lacking striae. Hind wing less than half length of abdomen. Terminalia (Fig. 7) with tergite IX not observed; sternite VIII (spiculum ventrale) extremely weakly lobed medially; proctiger narrowly triangular; paraprocts short, with ventral baculi converging apically; gonocoxites slightly shorter than paraprocts, each coxite with proximal plate more heavily sclerotized; gonostyli apparently absent. Distribution. Primarily along the drier slopes and valleys of the Coast Ranges in southern California but extending at least to Owens Valley (Inyo Co.) to the northwest and south into Baja California (Fig. 8). Note on Variation and Type Material. LeConte (1859a) did not mention how many specimens his original description was based on. Four male specimens are deposited in the LeConte collection (MCZ), three of which bear a gold circle (= “ California ”); the latter should probably be considered syntypes. The examined specimens of A. piceus demonstrate less individual and inter-populational variation than is seen in A. parvicollis. Specimens of the former from California and Baja California tend to be similar in terms of coloration and sclerotization. Specimens from the northeastern extent of its range in Nevada tend to be slightly smaller and lighter in color, and the female elytra are proportionately shorter than those of the single female known from California. Specimens Examined (671). UNITED STATES: CALIFORNIA: Inyo Co.: 7 miles west of Westgard Pass, 25 June 1953, H. Washburn (1♂, LACM); Alabama Hills, 36.60083°, 118.10615°, 1,350 m, 1 June 2016, UV/MV light, M. and L. Gimmel, A. Mayor, M. A. Johnston, C. Wirth, ML83 (1♂, SBMNH); Antelope Spring, 8 mi SW of Deep Springs, 29 June 1961, J. A. Litsinger (1♂, MAIC); same except 9 July 1962 (1♂, MAIC); Argus Mountains, 8 June 1938, J. A. Comstock (6♂♂, LACM); Baker Creek Campground, Big Pine, 37° 10.180 N, 118° 18.550 W, 8 June 2011, M. Kippenhan (1♂, MAIC); Big Pine Canyon, 8,5000, 8 August 1973, G. R. Ballmer (2♂♂, UCRC); Independence, 6 June 1918, L. L. Muchmore (2♂♂, LACM); Independence, 1 June 1972 (4♂♂, UCRC); Junction Ranch, 28 June 1969, L. Lester (2♂♂, LACM); Wyman Canyon, White Mountains, 9 July 1967, S. and S. Fromer (3♂♂, UCRC); 5 mi W of Big Pine, Baker Creek, 37.15424°N, 118.34295°W, 4,939 ft, 17 May 2004, UV light and trap, Thomas E. Dimock, 1121 (4♂♂, SBMNH); Kern Co.: Democrat Hot Springs, 2,2000 elevation, 16 June 1969, D. Levin (1♂, LACM); Keene, 3,000 ft, 7 July 1975, A. Fuller (1♂, SBMNH); Rancheria Creek in Piute Mountains, 4,3500 elevation, T29S R33E S23, 1 June 1973, J. P. and K. E. Donahue (2♂♂, LACM); Los Angeles Co.: 1 mile south of Valyermo, 19 May 1964, M. J. Wargo (3♂♂, UCRC); 2.5 miles SSW of Valyermo, 20 June 1959, N. McFarland (5♂♂, LACM); 5 miles north of Beverly Hills in Santa Monica Mountains, 1,1000 elevation, 13 July 1955, N. McFarland (3♂♂, LACM); Albright (2♂♂, LACM); Altadena, 10 May 1959, J. R. Northern (1♂, LACM); Angeles National Forest, San Dimas Experimental Forest, Bell Canyon, 34.1826°N, 117.7938°W, 13 June 2007, M. Caterino and S. Chatzimanolis (5♂♂, SBMNH); Angeles National Forest, Ruby Canyon, 34.6060°N, 118.5523°W, 22 June 2007, at light, M. Caterino and S. Chatzimanolis (4♂♂, SBMNH); Angeles National Forest, San Dimas Experimental Forest, Big Dalton Canyon, 34.1793°N, 117.7985°W, 27 July 2007, M. Caterino (1♂, SBMNH); Angeles National Forest, Shake Canyon, 34.7009°N, 118.5250°W, 2 July 2007, at light, M. Caterino (1♂, SBMNH); Big Pine Ranger Station, 5 July 1975, A. V. Evans (16♂♂, LACM); Big Pines, 13 July 1971 (6♂♂, LACM); Big Pines Recreation Area, 7 July 1983, S. Ziff (3♂♂, LACM); Big Pines Recreation Area, 20 July 1983, S. Ziff (2♂♂, LACM); Bouquet Canyon, 23 June 1937, N. Westerlund (3♂♂, LACM); Bouquet Canyon, 26 June 1937, J. A. Comstock (9♂♂, LACM); Bouquet Canyon, 7 July 1937, N. Westerlund and L. M. Martin (16♂♂, LACM); Burbank, 21 May 1930, C. H. Hicks (1♂, CASC); Burbank, 23 May 1930, C. H. Hicks and F. C. Hadden (3♂♂, LACM); Burbank, 30 June 1958 (2♂♂, LACM); Chatsworth, 26 June 1920, L. L. Muchmore (2♂♂, LACM); Claremont, 19 May 1962, T. Reimo (2♂♂, UCRC); Devil’ s Punch Bowl, 21 June 1964, D. G. Marqua and P. H. Sullivan (1♂, LACM); Devil’ s Punchbowl, 11 June 1975 (15♂♂, LACM); Eaton Canyon Park, 5 June 1964, R. H. Crandall (2♂♂, LACM); Glendale, 3 July 1965, E. I. Schlinger (1♂, UCRC); Gorman, 22 July 1938, J. von Bloeker (6♂♂, LACM); Juniper Hills, 8 June 1975, A. V. Evans (23♂♂, LACM); Juniper Hills, 4,3000 elevation, 14 July 1979, C. L. Bellamy (2♂♂, LACM); La Canada, 8 July 1950, D. C. Blodget (3♂♂, LACM); La Canada, 21 June 1969 (1♂, UCRC); La Crescenta, 19 June 1955, J. Roberds (6♂♂, LACM); La Verne, 24 June 1961, R. S. Erdman (4♂♂, LACM); Lake Hughes, 13 July 1938 (1♂, LACM); Littlerock, [day/month blank] 1945 (1♂, LACM); Littlerock, [day blank] May 1947, G. P. Mackenzie (1♂, LACM); Mint Canyon, 21 June 1949, H. E. Cott (1♂, SBMNH); Mount Wilson (2♂♂, LACM); Pomona, 6 April 1931, W. D. Clancy (1♂, UCRC); Santa Monica, 22 July 1943, G. P. Mackenzie (1♂, UCRC); Sierra Madre, 11 July 1944 (1♂, LACM); Sunland, 12 May 1940 (1♂, LACM); Tanbark Flats, 4 July 1950, D. C. Blodget (1♂, LACM); Tujunga, 19 June, 1955, J. Roberts (1♂, LACM); Tujunga, 14 May 1969, J. C. von Bloeker (30♂♂, LACM); Tujunga Canyon, 15 May 1969, A. Strong (1♂, UCRC); Littlerock, 8 June 1941, G. P. Mackenzie (1♂, UCRC); Topanga Canyon, 6 June 1978 (1♂, LACM); Pasadena, 25 May 1894 (1♂, LACM); Pasadena, [day blank] May [year blank] (1♂, LACM); San Marino, 5 July 1942, G. P. Mackenzie (1♂, UCRC); Valyermo, 1 July 1967, S. Ziff (2♂♂, LACM); West Los Angeles, 17 March 1967, B. S. Cheary (1♂, UCRC); [county record only], [day

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