Sclerocoelus copiosus Kuwahara, Marshall & Paiero, 2025, sp. nov.

Sclerocoelus copiosus sp. nov. urn:lsid:zoobank.org:act: 01A777A8-0CFC-46B5-90F6-D739DF37B0D2 Figs 1B, 5J, 34–36 Etymology The species name (from the Latin ‘ copiosus ’, meaning ‘copious, abundant’) reflects the abundance of this very widespread Neotropical species, especially in Malaise trap and flight-intercept trap samples from Costa Rica. Approximately 24% of the Neotropical Sclerocoelus specimens in the University of Guelph Insect Collection belong to this species. Material examined Holotype COSTA RICA • ♂; Puntarenas, Monteverde Reserve; 1500 m a.s.l.; 15–20 Aug.1986; L. Masner leg.; MNCR. Paratypes COSTA RICA – Alajuela • 5 ♂♂, 5 ♀♀; Quesada, Albergue Pozo Verde; 1800 m a.s.l.; 10°15′15.1″ N, 84°22′18.4″ W; 21 Apr. 2023; S.A. Marshall leg.; from refuse pile under Eciton burchellii bivouac; DEBU • 5 ♂♂, 6 ♀♀; same data as for preceding; MNCR • 2 ♂♂, 1 ♀; same data as for preceding; W. Porras leg.; MNCR. – Cartago • 1 ♂; Paraiso, Tapantí National Park; 9°43′21″ N, 83°46′30″ W; 1600 m a.s.l.; 28 Jul.–4 Aug. 2013; Proyecto ZADBI leg.; Malaise trap; MNCR • 1 ♂, 1 ♀; Tapantí National Park; 1650–1750 m a.s.l.; 7 Oct.1999; S.A. Marshall leg.; trail sweep; MNCR • 2 ♀♀; Tapantí National Park, above Ranger Station; 1250 m a.s.l.; 12 Oct. 1999; S.A. Marshall leg.; trail/road sweep; DEBU • 2 ♀♀; Tapantí National Park, Arboles Caídos trail; 1300 m a.s.l.; 9–11 Oct. 1999; S.A. Marshall leg.; sweep; MNCR • 7 ♂♂; Tapantí National Park, near west entrance, trail to hydro tower/ creek, 1150 m a.s.l.; 11 Oct. 1999; Marshall and Buck leg.; sweep; DEBU • 10 ♂♂; same data as for preceding; MNCR • 28 ♂♂, 15 ♀♀; Tapantí National Park, outside near west entrance; 1150 m a.s.l.; 8–9 Oct.1999; Marshall and Buck leg.; trail, sweep, DEBU • 30 ♂♂, 18 ♀♀; same data as for preceding; MNCR • 3 ♀♀; Tapantí National Park, Ranger Station; 1200 m a.s.l.; 7–11 Oct.1999; M. Buck leg.; pans in kitchen refuse; DEBU. – Puntarenas • 3 ♂♂, 4 ♀♀; same data as for holotype; MNCR • 6 ♂♂; Las Alturas Biological Station; 8°57′ N, 82°58′ W; 1500–1700 m a.s.l.; 12–14 Aug. 1995; S.A. Marshall leg.; DEBU • 6 ♂♂; same data as for preceding; MNCR • 2 ♂♂, 1 ♀; Monteverde Biological Reserve; 1500 m a.s.l.; 14 Jun. 2000; M. Buck leg.; sweeping treefall and trail; MNCR • 16 ♂♂, 10 ♀♀; Monteverde Biological Reserve; 1500 m a.s.l.; 14 Jun. 2000; M. Buck leg.; treefall, sweep and pans; DEBU • 16 ♂♂, 15 ♀♀; same data as for preceding; MNCR. – San José • 7 ♂♂, 5 ♀♀; Moravia, Zurquí de Moravia, Creek 2 north; 10°02′58″ N, 84°00′57″ W; 1600 m a.s.l.; 12–19 Apr. 2013; Proyecto ZADBI leg.; Malaise trap #2; MNCR • 5 ♂♂, 1 ♀; same data as for preceding; 19–26 Apr. 2013; MNCR • 1 ♂; same data as for preceding; 13–20 Sep. 2013; MNCR • 1 ♂; Moravia, Zurquí de Moravia, Tower path; 10°02′58″ N, 84°00′57″ W; 1600 m a.s.l.; 28 Feb.–8 Mar. 2013; Proyecto ZADBI leg.; Malaise trap #1, 0 m.; MNCR • 1 ♂; same data as for preceding; 26 Apr.–3 May 2013; MNCR • 2 ♂♂, 4 ♀♀; same data as for preceding; 30 Aug.–6 Sep. 2013; MNCR • 1 ♀; same data as for preceding; 11–18 Oct. 2013; MNCR. Other material examined BOLIVIA – La Paz • 20 ♂♂, 18 ♀♀; 0.5 km SE of Coroico; 16°11′ S, 67°44′ W; 15 Apr. 2001; S.A. Marshall leg.; DEBU • 1 ♂; 10 km NW of Caranavi, road to ENTEL tower; 15°46′35″ S, 67°35′48″ W; 1400 m a.s.l.; 13 Apr. 2001; S.A. Marshall leg.; DEBU • 7 ♂♂, 5 ♀♀; 23 km NE of Caranavi; 14 Apr. 2001; S.A. Marshall leg.; roadside/stream, sweep; DEBU • 12 ♂♂, 1 ♀; Chulumani, Apa-Apa Reserve; 16°21′15″ S, 67°30′21″ W; 2000 m a.s.l.; 1 Apr. 2001; S.A. Marshall leg.; sweep; DEBU. COSTA RICA – Alajuela • 1 ♂; La Virgen del Socorro, near Carriblanca; 700 m a.s.l.; 16 Feb. 1989; Grimaldi and DeVries leg.; AMNH • 1 ♂, 2 ♀♀; Río Peñas Blancas; 700 m a.s.l.; 18 Jul. 1986; L. Masner leg.; CNCI • 1 ♀; Volcán Tenorio, Bijagua Biological Station; 700 m a.s.l.; 20 Jun. 2000; M. Buck leg.; cut wet field, pan traps; DEBU. – Cartago • 3 ♂♂, 2 ♀♀; Braulio Carrillo National Park; 500 m a.s.l.; 10 Apr. 1985; L. Masner leg.; rainforest; CNCI • 63 ♂♂, 52 ♀♀; Braulio Carrillo National Park; 1400–1500 m a.s.l.; 11 Apr. 1985; H. Goulet and L. Masner leg.; cool moist river bed, Selva Premontana; CNCI • 1 ♂, 1 ♀; Highway 2, km 67; 6 Aug. 1995; S.A. Marshall leg.; roadside; DEBU • 2 ♂♂; Río Grande de Orosí, near Tapantí National Park; 1100–1150 m a.s.l.; 9 Oct. 1999; Marshall and Buck leg.; floodplain and forest; DEBU • 1 ♂, 4 ♀♀; same data as for preceding; 10–11 Oct. 1999; bromeliad leaf pile, sweeping and pans; DEBU • 22 ♂♂, 10 ♀♀; Río Macho Forest Reserve; 2200 m a.s.l.; 10 Oct. 1999; S.A. Marshall leg.; sweeping; DEBU • 1 ♂, 3 ♀♀; Tapantí National Park; 1550 m a.s.l.; 7–12 Oct. 1999; Marshall and Buck leg.; pans in fallen tree; DEBU • 13 ♂♂, 3 ♀♀; Tapantí National Park; 1650 m a.s.l.; 7 Oct. 1999; S.A. Marshall leg.; trail sweep; DEBU • 46 ♂♂, 20 ♀♀; Tapantí National Park; 1650–1750 m a.s.l.; 7 Oct. 1999; M. Buck leg.; sweep trail; DEBU • 37 ♂♂, 21 ♀♀; same data as for preceding; S.A. Marshall leg.; sweep trail; DEBU • 33 ♂♂; Tapantí National Park, above Ranger Station; 1250 m a.s.l.; 12 Oct. 1999; S.A. Marshall leg.; sweep trail/road; DEBU • 1 ♂; Tapantí National Park, Arboles Caídos trail; 1200 m a.s.l.; 4 Oct. 1999; Marshall and Buck leg.; trail sweep; DEBU • 3 ♂♂; same data as for preceding; 1300 m a.s.l.; 9/ 11 Oct. 1999; S.A. Marshall leg.; sweep; DEBU • 1 ♂, 1 ♀; Tapantí National Park, near Catarata trail; 1450 m a.s.l.; 4 Oct. 1999; S.A. Marshall leg.; roadside, sweeping leaf litter; DEBU • 97 ♂♂, 34 ♀♀; Tapantí National Park, near west entrance, trail to hydro tower/creek; 1150 m a.s.l.; 11 Oct. 1999; Marshall and Buck leg.; sweeping; DEBU • 7 ♂♂, 2 ♀♀; Tapantí National Park, outside near west entrance; 1150 m a.s.l.; 8–9 Oct. 1999; Marshall and Buck leg.; sweep trail; DEBU • 2 ♂♂, 1 ♀; Tapantí National Park, Ranger Station; 1200 m a.s.l.; 9–12 Oct. 1999; Buck and Marshall leg.; human dung, hand and traps; DEBU • 4 ♂♂; same data as for preceding; 7–11 Oct. 1999; M. Buck leg.; pans in kitchen refuse; DEBU • 1 ♂, 2 ♀♀; Tapantí National Park, road near represa [dam]; 1650 m a.s.l.; 8 Oct. 1999; M. Buck leg.; fallen tree; DEBU • 1 ♀; Tapantí National Park, trail to hydro tower near west entrance; 1150 m a.s.l.; 9–12 Oct. 1999; M. Buck leg.; pan traps; DEBU. – Heredia • 3 ♀♀; 6 km ENE of Vara Blanca; 10°11′ N, 84°07′ W; 2000 m a.s.l.; 13 Feb. 2002; DEBU • 4 ♂♂, 3 ♀♀; same data as for preceding; 21 Feb. 2002; DEBU • 4 ♂♂, 2 ♀♀; same data as for preceding; 20 Mar. 2002; DEBU • 6 ♂♂, 1 ♀; 16 km SSE of La Virgen; 10°16′ N, 84°05′ W; 1050–1150 m a.s.l.; 11–20 Feb. 2001; transect; DEBU • 49 ♂♂, 79 ♀♀; Moravia, near border of Braulio Carrillo National Park; 3–4 Mar. 1996; L. Masner leg.; creek bed, yellow pans; DEBU • 1 ♀; Santo Domingo, INBio Park; 9°58′23″ N, 84°05′30″ W; 6–7 Mar. 1996; L. Masner leg.; urban, yellow pans; DEBU. – Puntarenas • 119 ♂♂, 257 ♀♀; same data as for holotype; CNCI • 9 ♂♂, 3 ♀♀; 2000 m a.s.l.; 12 Aug. 1995; S.A. Marshall leg.; DEBU • 4 ♀♀; Las Alturas Biological Station; 8°57′ N, 82°58′ W; 1500–1700 m a.s.l.; 12–14 Aug. 1995; S.A. Marshall leg.; DEBU • 8 ♂♂, 2 ♀♀; Las Alturas Biological Station; 1550 m a.s.l.; 17 Aug. 1995; T. Pape leg.; DEBU • 28 ♂♂, 13 ♀♀; Las Alturas; 1500 m a.s.l.; 12 Aug. 1995; S.A. Marshall leg.; sweep forest-pasture margin; DEBU • 8 ♂♂, 2 ♀♀; same data as for preceding; 1500–1700 m a.s.l.; 12 Aug. 1995; sweep; DEBU • 5 ♂♂, 6 ♀♀; same data as for preceding; 1600 m a.s.l.; 11–14 Aug. 1995; Malaise trap; DEBU • 7 ♂♂, 3 ♀♀; same data as for preceding; 14 Aug. 1995; DEBU • 6 ♂♂, 11 ♀♀; same data as for preceding; 15 Aug. 1995; ground Eciton raid; DEBU • 1 ♂; same data as for preceding; swept over dung; DEBU • 1 ♂, 1 ♀; same data as for preceding; 2000 m a.s.l.; 15 Aug. 1995; treefall; DEBU • 1 ♀; Las Alturas, trail to Cerro Chai; 1700–2100 m a.s.l.; 13–15 Aug. 1995; S.A. Marshall leg.; DEBU • 1 ♀; Las Alturas, summit of Cerro Chai; 2100 m a.s.l.; 11–17 Aug. 1995; T. Pape leg.; DEBU • 23 ♂♂, 16 ♀♀; Monteverde Biological Reserve; 1500 m a.s.l.; 13 Jun. 2000; S.A. Marshall leg.; DEBU • 91 ♂♂, 52 ♀♀; same data as for preceding; 11–13 Jun. 2000; cloud forest; DEBU • 1 ♂, 1 ♀; same data as for preceding; 12–13 Jun. 2000; M. Buck leg.; pans along stream; DEBU • 3 ♂♂, 1 ♀; same data as for preceding; 13–14 Jun. 2000; DEBU • 85 ♂♂, 59 ♀♀; same data as for preceding; 11 Jun. 2000; sweep; DEBU • 1 ♀; same data as for preceding; on log over trail; DEBU • 5 ♂♂, 1 ♀; same data as for preceding; 12 Jun. 2000, S.A. Marshall leg.; on dung; DEBU • 9 ♀♀, 5 ♂♂; same data as for preceding; 26 May 1998; sweep; DEBU • 11 ♂♂, 33 ♀♀; same data as for preceding; 14 Jun. 2000; M. Buck leg.; sweeping treefall and trail; DEBU • 56 ♂♂, 67 ♀♀; same data as for preceding; treefall, sweep and pans; DEBU • 13 ♂♂, 9 ♀♀; same data as for preceding; 18 Mar. 1996; L. Masner leg.; along creek, yellow pans; DEBU • 2 ♂♂, 5 ♀♀; same data as for preceding; 12 Jun. 2000; M. Buck leg.; white pans in kitchen compost; DEBU • 1 ♂, 6 ♀♀; same data as for preceding; 13 Jun. 2000; DEBU • 2 ♂♂, 1 ♀; Monteverde Biological Station, lower trail; 26 May 1998; S.A. Marshall leg.; DEBU • 28 ♂♂, 88 ♀♀; Monteverde Reserve; 1500 m a.s.l.; Feb. 1980; W. Mason leg.; cloud forest; CNCI • 11 ♂♂, 26 ♀♀; Monteverde Reserve; 1550 m a.s.l.; 15–20 Jul. 1986; L. Masner leg.; CNCI • 1 ♂, 1 ♀; Monteverde; 10°18′ N, 84°48′ W; 1539 m a.s.l.; 24 Feb.–2 Mar. 1988; B. Hubley leg.; 1° tropical cloud forest, pan traps; ROME • 1 ♂, 1 ♀; Monteverde; 1500 m a.s.l.; 11–18 Jul. 1983; D. Lindeman leg.; fruit pitfall; CNCI • 2 ♂♂; Monteverde, 1500 m a.s.l.; 20 Feb. 1980; D.M. Wood; CNCI • 1 ♂; Monteverde; 1500 m a.s.l.; 24 Feb. 1991; B.J. Sinclair leg.; DEBU • 2 ♂♂, 9 ♀♀; Monteverde; 1500 m a.s.l.; 29 Feb. 1980; W.R. Mason leg.; cloud forest; CNCI • 3 ♂♂, 2 ♀♀; Monteverde; 1500 m a.s.l.; 4–11 Feb. 1989; D. Grimaldi leg.; lower montane wet forest; AMNH • 1 ♂, 4 ♀♀; Monteverde; 1500 m a.s.l.; 29 Feb. 1980; Mason and Wood leg.; rainforest; CNCI • 1 ♂, 1 ♀; Montev

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