Secular changes in colony‐forms and bryozoan carbonate sediments through geological history
Bibliographic record
Abstract
Abstract Ever since their first radiation in the Ordovician, bryozoans have contributed significantly to carbonate sedimentation. Most of the numerous colony‐forms developed by bryozoans have evolved repeatedly in different taxonomic groups and vary in their sediment‐producing potential. There are nine basic bryozoan colony‐forms: encrusting, dome‐shaped, palmate, foliose, fenestrate, robust branching, delicate branching, articulated and free‐living. The proportion of these morphotypes in bryozoan faunas period by period is shown to change significantly through the Phanerozoic. Notable patterns include: (i) steady increase in the number and proportion of encrusting species through time, interrupted by a transient drop in the Late Palaeozoic; (ii) post‐Triassic decrease in robust branching colonies; (iii) rise in the proportion of fenestrate colonies through the Palaeozoic, followed by their absence in the Triassic and Jurassic, rarity in the Cretaceous and reappearance in smaller proportions in the Cenozoic; and (iv) scarcity of articulated colonies and absence of free‐living colonies until the Cretaceous. Most Palaeozoic bryozoan sediments come from two architecturally distinct groups of colonies: (i) domal, delicate branching, robust branching and palmate; and (ii) fenestrate. The former generate coarse particles both as sediment and components of stromatoporoid‐coral reefs in the Early and mid Palaeozoic, whereas the delicate lacy fans of the latter create both prolific coarse sediment and form the cores of Late Palaeozoic deep‐water, sub‐photic biogenic mounds. Nearly all post‐Palaeozoic bryozoan sediments comprise cyclostomes and cheilostomes with many of the same growth forms but with the addition of free‐living colonies and significant numbers of articulated colonies. The latter produced sand and mud‐sized bryozoan sediment via disarticulation for the first time. In contrast to the Palaeozoic, post‐Palaeozoic bryozoans generated sediment varying more widely across the grain‐size spectrum, from mud to sand to gravel. This article highlights the need to consider evolutionary changes in carbonate‐producing organisms when interpreting facies changes through time.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.002 | 0.001 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.003 | 0.001 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".