Small‐Subunit rRNA Phylogenies Suggest That<i>Epalxella antiquorum</i>(Penard, 1922) Corliss, 1960 (Ciliophora, Odontostomatida) Is a Member of the Plagyopylea
Bibliographic record
Abstract
The odontostomatid ciliates have remained a homogeneous order of ciliates since the 1930s when they were recognized as a monophyletic assemblage. Since that time they have been placed with the heterotrich ciliates, and more recently transferred as incertae sedis to the new "riboclass" class Armophorea. We were able to obtain the small subunit rRNA gene sequence of the odontostomatid Epalxella antiquorum (Penard, 1922) Corliss, 1960, collected from the meromictic alpine Lake Alat in Germany, in July 2005. An alignment with representatives of all 11 classes of ciliates unambiguously places the Epalxella sequence with other representatives of the class Plagiopylea with 100% support in both maximum likelihood and Bayesian analyses. Epalxella is the basal lineage with trimyemid and plagiopylid ciliates forming the two terminal sister clades. While this molecular support is strong and unambiguous, there are no obvious morphological features to unite these three clades. Thus, the class Plagiopylea must continue to be referred to as a "riboclass." Using the Epalxella sequence as a basal marker, we tentatively identified 20 environmental sequences to the terminal plagiopylean clades: eight to the genus Trimyema; four to the genus Plagiopyla; and eight to two new species, one of which might represent a new plagiopylean genus.
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How this classification was reachedexpand
Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.001 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.001 | 0.000 |
| Bibliometrics | 0.000 | 0.000 |
| Science and technology studies | 0.000 | 0.001 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.001 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.000 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one teacher head, not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".