Phylogenetic relationships in the tribes Schizopetaleae and Thelypodieae (Brassicaceae) based on nuclear ribosomal ITS region and plastid<i>ndh</i>F DNA sequences
Bibliographic record
Abstract
Sequence data from the nuclear ribosomal internal transcribed spacer (ITS) region and plastid gene ndhF of 95 species, represented by 147 accessions, were used to determine the tribal limits, monophyly status, and phylogenetic intra-tribal relationships of genera within the New World tribe Schizopetaleae (formerly Thelypodieae; Brassicaceae). Maximum parsimony, Bayesian, and maximum-likelihood analyses all support the separate recognition of three distinct and generally well-supported clades. The largest corresponds to the North–South American Thelypodieae (27 genera: Catadysia , Caulanthus , Chaunanthus , Chilocardamum , Chlorocrambe , Coelophragmus , Dictyophragmus , Dryopetalon , Englerocharis , Hesperidanthus , Mostacillastrum , Neuontobotrys , Polypsecadium , Pringlea , Pterygiosperma , Romanschulzia , Sibara , Sibaropsis , Stanleya , Streptanthella , Streptanthus , Thelypodiopsis , Thelypodium , Thysanocarpus , Warea , Weberbauera , and Werdermannia ). The remaining six South American Schizopetaleae genera were divided into two clades: SCHIZ I of four genera ( Aschersoniodoxa , Brayopsis , Eudema , and Onuris ) and SCHIZ II Schizopetaleae s. str.: Mathewsia , and Schizopetalon ). Resolution within the Thelypodieae clade was limited in both the ITS- and ndhF-based phylogenies. Based on ITS sequence data, elements of the Old World tribe Brassiceae were sister to the Thelypodieae, whereas the ndhF data strongly supported the Sisymbrieae as sister to the Thelypodieae, and the Brassiceae as sister to both tribes. Sister groups to the Schizopetaleae clades I and II were not clearly resolved. Morphological and cytological data support the separation of these three clades. In SCHIZ II, the sepals are always erect to form a closed tube, and the trichomes are mostly dendritic, although other types also occur. In both the Thelypodieae and SCHIZ I clades, the sepals are variously oriented but never form a closed tube, whereas the trichomes are either absent, or simple, forked, or rarely dendritic. The latter clade differs from both SCHIZ II and Thelypodieae by lacking trichomes or rarely having forked ones and by having exclusively basal, entire leaves. Thelypodieae circumscription should be restricted in future to the Thelypodieae clade, Schizopetaleae circumscription to the SCHIZ II clade, while a new tribe is needed for the SCHIZ I clade.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.001 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".