Description of Leptoomus janzeni, n. gen. and n. sp. (Hymenoptera: Chalcidoidea) from Baltic amber, and discussion of its relationships and classification relative to Eupelmidae, Tanaostigmatidae and Encyrtidae
Bibliographic record
Abstract
Leptoomus janzeni, n. gen. and n. sp., is described from 5 females and 2 males in 7 pieces of Baltic amber. An enlarged acropleuron forming the entire mesopleuron places the taxon within Eupelmidae, Tanaostigmatidae or Encyrtidae (Chalcidoidea), but it has a unique combination of features that differentiates it from extant members of these families. Its structural features are compared with those that characterize the three families and it is postulated to be the sister group of Tanaostigmatidae sensu stricto + (Cynipencyrtus + Encyrtidae) based on relative structure of its pronotum, prepectus and mesothoracic spiracle, and retention of two other putative symplesiomorphies, the presence of complete furrowlike notauli and a protibial apical spicule. Conflicting character states indicate other possible relationships, including a sistergroup relationship with Cynipencyrtus + Encyrtidae based on transverse-triangular axillae and a flagellum having only seven funicular segments, or with Tanaostigmatidae s. s. based on presence of an externally visible prepectal pouch, or possibly forming a monophyletic group with Tanaostigmatidae s. s. + Cynipencyrtus based on combined acropleuralmetacoxal structure. Several features support the monophyly of Tanaostigmatidae s. s., that is excluding Cynipencyrtus and Leptoomus, but these two genera are both classified in Tanaostigmatidae sensu lato until relationships are resolved more conclusively. Features possessed by different members of Neanastatinae (Eupelmidae) suggest that this subfamily may be closely related to Tanaostigmatidae s. l. + Encyrtidae, but possible relationships of Tanaostigmatidae s. s. are also discussed if its enlarged acropleuron and other skeletomusculature features associated with jumping are convergent to similar features in the other taxa.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.002 | 0.001 |
| Science and technology studies | 0.001 | 0.001 |
| Scholarly communication | 0.000 | 0.001 |
| Open science | 0.001 | 0.001 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.001 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".