Tidying up the genus<i>Letharia</i>: introducing<i>L. lupina</i>sp. nov. and a new circumscription for<i>L. columbiana</i>
Bibliographic record
Abstract
Abstract Western North America is the global centre of diversity forLetharia, a distinctive and cryptically diverse genus of lichenized fungi belonging to theParmeliaceae. The genus is characterized by a shrubby, fruticose habit and presence of vulpinic acid. Previous studies using multiple fungal nuclear loci revealed the existence of two distinct species-level lineages within the traditional concept ofL. vulpinaand four such lineages withinL. columbiana. Here we use molecular sequence data in an attempt to settle long-standing taxonomic issues in the genus. Our results confirm the widespread existence withinL. vulpinas. lat. of two distinct species-level groups, each forming a mutually exclusive partnership with a separate algal clade withinTrebouxia jamesiis. lat. Accordingly, we formally describe the segregate speciesL. lupinasp. nov. Our results also support the evolutionary independence of four candidate species previously circumscribed fromL. columbianas. lat. One of these lineages,L.‘gracilis’, has already received species recognition asL. gracilis, while a second,L.‘lucida’, is epitypified here againstL. columbianas. str. Based on results from species delimitation analyses under the multispecies coalescent model, the two remaining lineages,L.‘barbata’ andL.‘rugosa’, also warrant formal taxonomic recognition; however, we refrain from describing these species pending additional studies of diagnostic characters, ecological preference, and distributions.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.000 |
| Science and technology studies | 0.001 | 0.001 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.001 | 0.001 |
| Research integrity | 0.000 | 0.001 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".