A comprehensive species sampling sheds light on the molecular phylogenetics of Calothecinae (Poaceae, Pooideae, Poeae): Evidence for a new subtribe and multiple genera within the <i>Chascolytrum</i> clade
Why this work is in the frame
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Bibliographic record
Abstract
Abstract The circumscription of the grass subtribe Calothecinae has undergone several changes since its description. Currently, it comprises Chascolytrum and the recently described genera Laegaardia and Paramochloa . Here we evaluate the circumscription of Calothecinae and the recently proposed infrageneric classification of Chascolytrum based on a phylogeny with more comprehensive taxon and molecular marker sampling than in previous studies. We sampled all Calothecinae genera, all but one Chascolytrum species, two South American Trisetum s.l ., and representatives of the subtribes Agrostidinae, Brizinae, Echinopogoninae, Koeleriinae, Phalaridinae, and Torreyochloinae within Poaceae tribe Poeae. We performed Bayesian and Maximum Likelihood analyses of four plastid DNA regions ( atpF‐atpH , matK , rps16 intron, and trnL‐trnF ) and two nuclear ribosomal regions (ITS and ETS). Our results revealed that neither Calothecinae nor Chascolytrum is monophyletic, as currently recognized, because Trisetum brasiliense and Trisetum bulbosum are nested within Chascolytrum . We include these two Trisetum species in Calothecinae as incertae sedis . Laegaardia and Paramochloa form a clade that is sister to the Chascolytrum + Trisetum clade, and based on morphological characters, we transfer the former to the new subtribe Paramochloinae. Our Chascolytrum phylogeny is better resolved and supported than in previous studies, and based on these results, we divide Chascolytrum into nine genera, including two new ones: Boldrinia (gen. nov.), Calotheca , Chascolytrum , Erianthecium , Lombardochloa , Microbriza , Poidium , Rhombolytrum , and Rosengurttia (gen. nov.). We provide a key to Calothecinae genera, descriptions of the genera, nomenclatural information, and keys to species of each genus. In addition, six new combinations are proposed.
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Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.000 | 0.000 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.000 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from it