Cranial Anatomy of the Caseid Synapsid Cotylorhynchus romeri, a Large Terrestrial Herbivore From the Lower Permian of Oklahoma, U.S.A
Bibliographic record
Abstract
The detailed description of the cranial anatomy of Cotylorhynchus romeri, a very large caseid synapsid from the lower Permian Hennessey Formation of Oklahoma, uncovered several potential autapomorphies, including parietal-postorbital contact greatly reduced by broad anterior process of supratemporal; the postparietals are transversely broad and contact the supratemporals laterally; the quadratojugal has a pennant-like occipital process; the stapes has a short shaft and a ventral process that abuts against the palate; the bulbous marginal dentition narrows distally and carries three small denticles; the vomer has three large teeth along its medial edge; parasphenoidal dentition is present; and the surangular overlaps the posterodorsal tip of the dentary and excludes it from the coronoid eminence. Owing to lack of comparative cranial material in most large caseids, the evolutionary history of these autapomorphies remains ambiguous because they cannot be determined in the closest relatives of Cotylorhynchus romeri . Our description of the skull of Cotylorhynchus romeri is consonant with the hypothesis that this caseid was a high-fibre terrestrial herbivore. The recent hypothesis that Cotylorhynchus romeri was primarily aquatic was proposed on a paleobiological basis that ignored paleontological and taphonomic evidence from the Hennessey Formation. Autochthonous preservation of several articulated skeletons of Cotylorhynchus romeri in subaerially deposited sediments that also preserve “swarms” of an aestivating fossil amphibian ( Brachydectes ) indicate that this caseid was the largest tetrapod of a terrestrial fauna that lived in a monsoonal climate.
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How this classification was reachedexpand
Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.001 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.000 | 0.001 |
| Science and technology studies | 0.001 | 0.002 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.001 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one teacher head, not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".