Incipient speciation in aquatic snails in an arid-zone spring complex
Bibliographic record
Abstract
The gastropod genus Caldicochlea is endemic to the Dalhousie Springs in the arid north of South Australia. The genus contains two species, or species complexes, Caldicochlea harrisi and C. globosa that overlap morphologically but can be distinguished genetically, principally by allozymes at the Lap, Sordh-1 and Sordh-2 loci. C. harrisi is found in many of the approximately sixty active springs in the 70 km2 of the Dalhousie complex, but C. globosa is confined to the northern half where there are larger, hotter springs. The species co-occur in many springs in this area, generally with apparently none, or low levels of hybridization, as judged by the absence of heterozygotes at the Lap and Sordh-2 loci. Introgression is apparendy low and bidirectional. In two springs within the current ranges of both species, Lap genotypes occur in Hardy-Weinberg expected frequencies. Genotypes at other loci suggest that this is unlikely to be due to the independent evolution of Lap polymorphisms in these springs. Reproductive isolation may have broken down or may not have been established in these springs. For the latter to be so, reproductive isolation must have evolved through intra-spring interactions between the species, and independently in different springs. Isolation would not be fully established, if at all, until after secondary contact of their colonizing populations.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.000 |
| Science and technology studies | 0.001 | 0.001 |
| Scholarly communication | 0.001 | 0.000 |
| Open science | 0.000 | 0.001 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".