Myotis septentrionalis

FIG. 2. Dorsal, ventral, and lateral view of the skull and lateral view of the mandible of a male Myotis septentrionalis (Provincial Museum of Alberta, Z79.116.1, from Cadomin, Alberta).Greatest length of skull 15.3 mm.Photographs by G. Newlands.FIG. 1. Myotis septentrionalis at a roost hole.Photograph by M. B. Fenton. Myotis septentrionalis (Trouessart, 1897)Northern Long-eared Bat Vespertilio gryphus var.septentrionalis Trouessart, 1897:131.Type locality ''Halifax, Nova Scotia.''Myotis keenii septentrionalis Miller and Allen, 1928:105.Myotis septentrionalis van Zyll de Jong, 1979:993.First use of current name combination. CONTEXT AND CONTENT.Order Chiroptera, suborder Microchiroptera, family Vespertilionidae, subfamily Vespertilioninae, genus Myotis, subgenus Myotis (Findley, 1972).This species was presented as a subspecies of M. keenii in Mammalian Species No. 121 (Fitch and Shump, 1979).M. keenii and M. septentrionalis occupy nonoverlapping ranges.Thus, any reference to M. keenii (Keen's myotis or Keen's bat) occurring outside of the Pacific Northwest of North America refers to M. septentrionalis.No subspecies are recognized. DIAGNOSIS.Myotis septentrionalis (Fig. 1) can be distinguished from M. lucifugus by its longer ears and tragus and relatively longer tail.M. septentrionalis is similar in color to M. lucifugus, but M. lucifugus has a glossier pelage.Unlike, M. lucifugus, the maxillary toothrow of M. septentrionalis exceeds width across molars.M. septentrionalis also has a least interorbital width/ length of maxillary toothrow ratio Ͻ0.7, whereas that of M. lucifugus is Ͼ0.7 (van Zyll de Jong, 1985).M. septentrionalis can be distinguished from M. evotis by its darker pelage and paler membranes, which give M. septentrionalis an overall brown look.M. keenii usually possesses dark shoulder spots and a sparse scattering of hair along free end of uropatagium, characteristics that are not present in M. septentrionalis.Skull of M. keenii is distinguished from that of M. septentrionalis by a width of canine at cingulum of Ͻ0.76 mm.Both M. keenii and M. evotis tend to have longer ears and a narrower skull (van Zyll de Jong, 1979). GENERAL CHARACTERISTICS.Head and body length Ͻ50 mm, with total length up to 95 mm.Hind foot is generally Ͻ60% length of tibia.Pelage and membranes are light brown; membranes usually concolor with pelage.Ears are long and usually extend past the nose when pushed forward.Third, fourth, and fifth metacarpals are of about equal length.Skull is narrow with a relatively long rostrum.Females are generally larger and heavier than males (Caire et al., 1979;Williams and Findley, 1979).Body mass is 5-8 g.Mean (van Zyll de Jong, 1985) and range Glass and Ward, 1959;Turner, 1974) of some external and skull measurements (in mm) are as follows (Fig. 2): length of forearm, 36.4 (34-38); length of ear, 16.4 (14-19); total length, 86.2 (77-95); length of tail, 37.7 (35-42); length of hind foot, 9.4 (8-10); greatest length of skull, 14.8 (14.6-15.6);mastoid breadth, 7.8 (7.4-7.9);length of maxillary toothrow, 5.8 (5.4-6.0);least interorbital width, 3.7 (3.4-3.8); and zygomatic breadth, 8.2-9.7.Other mean cranial measurements (in mm) include cranial depth, 5.0

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