Phylogeny, Classification, and Character Evolution of <i>Acalypha</i> (Euphorbiaceae: Acalyphoideae)
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Bibliographic record
Abstract
Abstract— Acalypha (Euphorbiaceae: Acalyphoideae) is a large, monophyletic genus distributed worldwide in tropical and subtropical regions, with a few species extending into temperate areas of southern Africa, Asia, and North and South America. We reconstructed phylogenetic relationships within the genus using DNA sequences from the plastid ndhF and trnL-F regions and the nuclear ribosomal ITS region, sampling 142 species to represent the geographic, morphologic, and taxonomic diversity with the genus, resulting in a 162 (158 in Acalypha ) terminal and 3847 character combined dataset. Bayesian and maximum likelihood reconstructions based on the combined dataset yielded a tree with a generally well-supported backbone and several strongly supported clades. Our results strongly supported the monophyly of Acalypha subg. Acalypha as currently recognized but showed that A. subg. Linostachys and almost all other infrageneric taxa recognized in the most recent comprehensive classification of the genus were not monophyletic. We therefore propose a new subgeneric classification comprising A. subg. Acalypha , A. subg. Androcephala , A. subg. Hypandrae , and A. subg. Linostachys (s.s.). Our results also shed light on relationships within some species groups, including in what has been treated as a broadly defined A. amentacea , in which we recognize A. amentacea , A. palauensis comb. nov. , and A. wilkesiana as distinct species. Bayesian ancestral state estimations based on the phylogeny of Acalypha demonstrated that inflorescence position and sexuality and habit show high homoplasy, especially within A. subg. Acalypha , and that inflorescence position and habit exhibit correlated evolution.
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Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.001 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.000 | 0.000 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.000 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from it