Empis (Enoplempis) scotti Sinclair, Brooks & Cumming, 2025, sp. nov.

Empis (Enoplempis) scotti sp. nov. urn:lsid:zoobank.org:act: 201C66EF-F42C-4651-9987-4BAA28E84D6F (Figs 207, 221–223, 237) Type material. HOLOTYPE ♂ (Figs 221–223), labelled: “USA: OR: Benton Co., Corvallis,/ Lewisburg Saddle, below Old / Growth Trail, 44°38′32.28″N / 123°17′20.76″W, Malaise trap,/ over creek, 23.iv–6.v.2013, S. Fitzgerald, CNC350416 ”; “ HOLOTYPE / Empis (Enoplempis) / scotti Sinclair,/ Brooks & Cumming [red label]” (CNC). PARATYPES: USA. California: Humboldt County: 2 mi. W Briceland [40°06′N 123°54′W], 21.v.1976, R. Dietz, J. Hafernik (1♂, EMEC). Mendocino County: Calif. Coast Range Pres., 5 mi. N Branscomb [39°42′N 123°39′W], 26.v.1976, JAP (1♂, EMEC); same data except, E. Rogers (1♂, EMEC); NCCRP, 3 mi. N Branscomb, 1400 ft, 17.v.1975, JAP (1♂, EMEC); same data except, 5–6.v.1989, MT, S. O’Keefe (1♂, EMEC). Oregon: Benton County: Corvallis, 3430 NW Deer Run St., 44°36′6″N 123°18′51″W, 20.iv–6.vi.2018, MT, SJF (1♂, 3♀, CNC); McDonald Forest, Oak Creek, 44.605623°N 123.332986°W, 20.v.2014, hillside woods and seep, SJF (1♂, CNC); Rock Ck, 5 mi. W Philomath [44.5020°N 123.4426°W], 2.v.1952, V. Roth (1♂, OSAC). Coos County: ca 3 mi. SE of Remote, Rock Ck, 42°59′27″N 123°51′41″W, 28.v.2009, SEB (1♂, CNC). Douglas County: ca 5.5 mi. SW Camas Valley, Twelvemile Ck, 42°58′41″N 123°44′29″W, 27.v.2009, J.M. Cumming (1♂, CNC); 7 mi. SW Camas Valley, Bear Ck Recreation Site, 42°58′08″N 123°45′55″W, 27.v.2009, SEB (1♂, CNC); 7 mi. SW Camas Valley, Bridge Ck at Twelvemile Ck Rd, 42°58′07″N 123°43′29″W, 27.v.2009, SEB (1♂, CNC). Washington County: Forest Grove [45°31′N 123°06′W], 12.iv.1919 (1♂, EMEC). Diagnosis. This species is distinguished from other species of the E. (En.) teres species group by the pair of dark scutal vittae, male hind femur strongly bowed on apical third with anteroventral row of strong setae, without tuft of whip-like setae at base of ventral process (Figs 222, 223), epandrium with long apical and ventral setae, and dorsal margin of male cercus straight. Description. Wing length 4.4–5.5 mm. Male. Similar to E. (En.) teres, except as follows: head narrowly dichoptic, frons below ocellar triangle slightly narrower than width of anterior ocellus; with short setulae along inner margin of eye. Occiput bearing single row of strong setae, shorter than ocellar setae; postocular setae shorter than occipital setae. Antenna with postpedicel long and tapered, 6× longer than basal width. Thorax with postpronotal lobe with pale shiny apex; postalar ridge yellowish. Scutum with 1–2 posterior npl. Scutellum with 1 pair of long sctl, with 1–3 pairs of marginal setulae. Legs slender, orange brown, including coxae, apex of tibiae darker, tarsi brown; coxae with greyish pruinescence (Fig. 221). Fore femur without white ventral pile. Fore tibia with several longer anterodorsal and posterodorsal setae on apical third, no longer than width of tibia; anterior face with short appressed setae; apex with long posterior and ventral preapical setae. Fore tarsomere 1 swollen, nearly subequal in width to width of apex of tibia; without spine-like ventral setae; tarsomeres 1–3 with long dorsal setae, longer than width of tarsomere; tarsomeres 2–5 with biserial row of spine-like ventral setae; tarsomeres 1–4 with apicolateral pair of spine-like setae, longer than width of segment. Mid femur with white ventral pile; row of short posteroventral setae, shorter that femur width. Mid tibia with anteroventral and posteroventral setae, shorter than width of tibia; rows of short anterodorsal and posterodorsal setae, similar to ventral setae. Mid tarsus with all tarsomeres with biserial row of spine-like ventral setae; tarsomeres 1–4 with apicolateral pair of spine-like setae, longer than width of segment. Hind femur with white ventral pile and anteroventral and posteroventral row of slender setae on basal two-thirds; apical third strongly bowed with anteroventral row of strong setae, shorter than width of femur; preapex with broad ventral expansion tapered to narrow posteroventral apex and flattened anteroventral hooked lobe; posteroventral face with row of 3 peg-like setae and upper row of long slender setae; apex without preapical row of posterior setae; strong preapical anterior seta and 2 preapical anterodorsal setae (Figs 222, 223). Hind tibia greatly narrowed and twisted at base; without anteroventral process; posteroventral process knob-like, with rounded apex crowned with dense, dark thickened setae and row of similar setae on anterior face of apex; dense row of thickened anteroventral setae extending from base of posteroventral process to near apex; long anterodorsal and posterodorsal setae, twice as long as width of tibia; anterior and ventral faces with similar shorter erect setae (Figs 221–223). Hind tarsus with long dorsal setae on tarsomeres 1–3; tarsomere 1 swollen, narrower than apex of tibia with strong ventral setae; tarsomeres 2–5 similar to fore tarsus. Halter yellowish to whitish. Abdomen paler than thorax, with dense whitish-grey pruinescence, with dark marginal setae. Terminalia (Fig. 207): brown with dorsal margin of cercus blackish; epandrial lamella with grey pruinescence. Cercus long, narrow, tapered to narrow apex; dorsal margin straight; dense setae along dorsal margin, shorter than width of cercus; base weakly fused to epandrial lamella. Subepandrial process extending on either side of phallus; apex narrow, downcurved. Epandrial lamella narrow, linear, longer than height; dorsal margin slightly expanded subapically; bearing long setae apically and ventrally, longer than width of lamella. Hypandrium very short, truncate, not prolonged laterally; without setae. Phallus with basal third narrow, tapered, gently arched anteriorly; apex emerging beyond cercus; apex slender and downcurved in lateral view; roughened, spiculate subapically, near dorsal margin of epandrium; ejaculatory apodeme elongate, more than half-length of epandrium, Y-shaped, with ventrally positioned lateral apodemes. Female. Similar to male, except as follows: frons broader, slightly wider than anterior ocellus ocellar triangle; scutum with 2 pairs of broader brownish vittae; hindleg without modified setae and processes. Abdominal pleural membrane not expanded; apex of tergite 8 shiny. Geographical distribution and seasonal occurrence (Fig. 237). Empis (En.) scotti sp. nov. is known from the Coast Range in Oregon and northern California. Adults have been collected in April and May. Etymology. This species is named after fellow dipterist Scott J. Fitzgerald, who collected the holotype. Nuptial gift presentation. Form unknown. Remarks. Empis (En.) fitzgeraldi sp. nov., E. (En.) undulasetosa sp. nov. and E. (En.) scotti sp. nov. are sympatric and were collected during the same collection event.

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